FOREST PHYSIOLOGY LAB
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Resident Researchers

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Dr. Lucy Kerhoulas
  • Associate Professor, Cal Poly Humboldt, Department of Forestry & Wildland Resources
  • Education
    • PhD, Biology, Northern Arizona University, 2013
    • MS, Biology, Humboldt State University, 2008
    • BS, Botany, Humboldt State University, 2006
  • Research Interests: tree physiology and water use, forest ecology, restoration, dendrochronology, stable isotopes, crown ecology, tree climbing
  • Chief Editor, Northwest Science
  • Courses taught
    • ​Dendrology (FOR 130)
    • ​Silvics (FOR 331)
  • Email: lucy.kerhoulas at humboldt dot edu
  • Curriculum vitae

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Dr. Nick Kerhoulas
  • Lab Manager and Lecturer, Cal Poly Humboldt, Department of Wildlife
  • Education
    • PhD, Biology, University of Alaska Fairbanks, 2017
    • MS, Biology, Humboldt State University, 2008
    • BS, Wildlife, Humboldt State University, 2006
  • Research Interests: phylogenetics, conservation biology, biogeography, small mammals, arboreal mammals, tree climbing
  • Courses taught
    • ​Conservation Biology (WLDF 460)
    • ​Mammals Management (WLDF 422)
    • Wildlife Techniques (WLDF 311)
  • Email: nicholas.kerhoulas at humboldt dot edu

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​Wade Polda
  • Lab Research Associate; helps with bird surveys, epiphyte identification, tree climbing, and field-based physiology measurements
  • Education: BS, Botany, Humboldt State University, 2016
  • Email: wtp1 at humboldt dot edu


MS Graduate Students

Current
  • Annie Allen — Grazing impacts on fen vegetation communities and their environmental drivers in the Klamath Mountains Ecoregion
  • Courtney Copper -- Influences of conifer encroachment on Oregon white oak crown structure and non-vascular epiphyte communities
  • David Dorval — Factors influencing tree vigor and structural failure along power line corridors in northern California
  • Sebastian Evans — Alpine plant succession in a recently deglaciated foreland basin in the Trinity Alps
  • Jenni Hedin — High elevation five-needle pines in the Klamath Mountains
  • Millie McCord — Ecophysiological effects of conifer removal treatments on Oregon white oaks in eastern Oregon
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Past
  • Zoë Ziegler — Patterns of tree damage and fuels around powerlines in northern California (2025) (PDF)
  • Gabriel Roletti — Drought responses across diverse conifer species, habitats, and competitive gradients in Northern California (2022) (PDF)
  • Sophia Lemmo — Drought, tree mortality, and regeneration in northern California (2022) (PDF)
  • Gabriel Goff — Conifer encroachment and removal in a Northern California oak woodland: Influences on ecosystem physiology and biodiversity (2021) (PDF)
  • Wallis Robinson — Influences on conifer drought responses in Northern California (2021) (PDF)
  • Kevin Soland — Efficacy of forest restoration across a 40-year chronosequence in Redwood National Park (2020) (PDF)
  • Ariel Weisgrau — Alternative water sources in western conifers via foliar uptake and hydraulic redistribution (2020) (PDF)


​Undergraduate Researchers

Current
  • Maddox Haley — High elevation five-needle pines in the Klamath Mountains
  • Gabriel Abundis — Effects of grazing on Klamath Mountain fens
  • Ryan Gilbert — Effects of grazing on Klamath Mountain fens
  • Heather Davis — Effects of grazing on Klamath Mountain fens
  • Finn Scott-Neff — Botanical shifts across time in the Klamath Mountains
  • Jack Christy — Effects of climate change and fire exclusion on overstory plants in the Klamath Mountains
  • Stella Stahl — Herbarium collections from the Klamath Mountains
  • Kaitlyn Briggs — Forest water resources in Northern California
  • ​Forrest Horobin — Klamath Mountains Ecoregion vegetation patterns
  • Corri Jai — Meadows monitoring in the Klamath Mountains

Past​
  • Redwoods Rising  — Hired 100 undergraduate students to work on redwood and giant sequoia restoration projects (2018 - 2025)
  • Wren Williams — Tree failure patterns in northern California, redwood and oak water sources (2024-2025)
  • Emily Jackson — Effects of climate change and fire exclusion on understory plants in the Klamath Mountains (2023-2024)
  • Maeve Flynn — Effects of climate change and fire exclusion on understory plants in the Klamath Mountains (2024)
  • Tori Bernal — EcoPlot data (2024-2025)
  • Hannah Ricketts — EcoPlot data (2024-2025)
  • Rain Lopez — Tree failure patterns in northern California (2024)
  • Damian Giles — Lab Manager (2023-2024)
  • Leyana Miller — Tree failure patterns in northern California (2024)
  • Daven Haywood — Tree failure patterns in northern California (2024)
  • Cameron Tavis — Restoration in Oregon white oak and California black oak woodlands (2024)
  • Olivia Fox — Restoration in Oregon white oak and California black oak woodlands (2024)
  • Mark Lara — Tree failure patterns in northern California (2024)
  • Sienna Tubridy — Tree failure patterns in northern California (2024)
  • Miya Tanner — Meadows monitoring in the Klamath Mountains (2024)
  • Grayson Prater — Effects of climate change and fire exclusion on montane understory plants in the Klamath Mountains (2024)
  • Jess Carey — Effects of climate change and fire exclusion on montane understory plants in the Klamath Mountains (2024)
  • Mouse Sloan — Effects of climate change and fire exclusion on montane overstory plants in the Klamath Mountains (2024)
  • Clarissa Peyton — Effects of climate change and fire exclusion on montane overstory plants in the Klamath Mountains (2024)
  • Sebastian Castillo — Wildfire influences on conifer drought responses (2024)
  • Ernesto Chavez-Velasco — Klamath Mountains Ecoregion vegetation patterns (2024)
  • Grayson Jameson — Tree failure patterns in northern California (2024)
  • Clare Dooley — Plant water sources in the redwood forest (2023)
  • Michelle Vongsa — Klamath Mountains Ecoregion vegetation patterns (2023)
  • Alec Brown — Klamath Mountains Ecoregion vegetation patterns (2023)
  • Fiona Shaw — Plant water sources in the redwood forest (2023)
  • Cody Ellis — Plant water sources in the redwood forest (2023)
  • Heather Davis — Plant water sources in the redwood forest (2023); Effects of climate change and fire exclusion on montane overstory plants in the Klamath Mountains (2024)
  • Dave Ramirez — Tree failure patterns in northern California (2023)
  • Cob Poss — Tree failure patterns in northern California (2023)
  • Hannah Dungan — Tree failure patterns in northern California (2023)
  • Lily Cummins — Black oak woodland restoration (2023)
  • ​Liam Hay — Black oak woodland restoration (2023)
  • Britney Martinez — Forest mortality and regeneration in northern California (2021)
  • Jenna Brown — Forest mortality and regeneration in northern California (2021)
  • Charles Russell — Forest mortality and regeneration in northern California (2021)
  • Sarah Aguiar — Conifer drought responses in northern California (2020)
  • Perris Alfonzo — Conifer drought responses in northern California (2020)
  • Ashley Shannon — Conifer drought responses in northern California (2020)
  • Sara Bandali — Conifer drought responses in northern California (2020)
  • Ian Conway — Conifer drought responses in northern California (2020)
  • Jeremy Dustin — Conifer drought responses in northern California (2020)
  • Maeve Flynn — Conifer drought responses in northern California (2020)
  • Rosalio Gonzalez — Forest mortality and regeneration in northern California (2020)
  • Suzanne Melendez — Conifer drought responses in northern California (2020)
  • Diane Orozco — Conifer drought responses in northern California (2020)
  • Ryan Maberry — Physiological effects of Ramorum leaf blight on understory hardwoods (2020)
  • Jessica Suoja — Physiological effects of Ramorum leaf blight on understory hardwoods (2020)
  • William Weinberg — Physiological effects of Ramorum leaf blight on understory hardwoods (2020)
  • Asher Budnik — Conifer drought responses in northern California (2019)
  • Elizabeth Hinojosa — Conifer drought responses in northern California (2019)
  • Brigitte Price — Conifer drought responses in northern California (2019)
  • Colleen Smith — Conifer drought responses in northern California (2019)
  • David Hammons — Foliar morphology and physiology in bigleaf maple (2019)
  • Meagan Burger — Redwood physiological responses to forest restoration (2018)​
  • Tyler Dunlap — Redwood physiological responses to forest restoration (2018)
  • Gabriel Goff — Physiological responses of Oregon white oak to conifer encroachment (2018)
  • Alex Gorman — Epiphyte diversity and distribution in an old Sitka spruce crown (2018)
  • Emily Hoeft — Foliar uptake in Sitka spruce (2018)
  • Sonnette Russell — Redwood physiological responses to forest restoration (2018)
  • Kevin Soland — Redwood physiological responses to forest restoration (2018)
  • Scott Whittington — Population assessment for the Sonoma tree vole at the L.W. Schatz Demonstration Tree Farm (2017)
  • Millen McCord — Epiphyte diversity in Sitka spruce crowns (2017)
  • Wade Polda — Physiology and growth of conifer seedlings planted after variable density retention treatments (2016)
  • Ariel Weisgrau — Adaptation to climate change? Moving coast redwood seedlings northward and inland (2015) & Foliar morphology and physiology of bigleaf maple (2016)
  • John Naccarato — Adaptation to climate change? Moving coast redwood seedlings northward and inland (2015)
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